Principal Product Designer - Customer Lifecycle - Ancestry

About Ancestry :

When you join Ancestry , you join a human-centered company where every person’s story is important. Ancestry ®, the global leader in family history, connects everyone with their past so they can discover, preserve, and share their unique family stories. With our unparalleled collection of more than 65 billion records, over 3.5 million subscribers, and over 27 million people in our growing DNA network, customers can discover their family story and gain a new level of understanding about their lives. Over the past 40 years, we’ve built trusted relationships with millions of people who have chosen us as the platform for discovering, preserving, and sharing the most important information about themselves and their families. We are committed to our location flexible work approach, allowing you to choose to work in the nearest office, from your home, or a hybrid of both (subject to location restrictions and roles that are required to be in the office- see the full list of eligible US locations HERE ). We will continue to hire and promote beyond the boundaries of our office locations, to enable broadened possibilities for employee diversity. Together, we work every day to foster a work environment that's inclusive as well as diverse, and where our people can be themselves. Every idea and perspective is valued so that our products and services reflect the global and diverse clients we serve. Ancestry encourages applications from minorities, women, the disabled, protected veterans and all other qualified applicants. Passionate about dedicating your work to enriching people’s lives? Join the curious.

We’re looking for a Principal Product Designer to join our End-to-End Experience (E2E) team — a specialized group that works across product areas to ensure our experiences feel cohesive, high-quality, and deeply valuable to customers. This role is a unique opportunity to shape how Ancestry shows up holistically, connecting touch-points into a seamless, intuitive journey.

This is not about owning a single product surface. It’s about seeing across them — identifying gaps, inconsistencies, and opportunities — and partnering with teams to elevate the entire ecosystem. You’ll also work closely with Product Marketing to ensure the value of our experiences is clear, compelling, and reflected consistently in the product.

You’ll report to the Design Director of the E2E team and collaborate closely with peers across product, design, engineering, research, content, marketing, and customer support. Your role is to help ensure that every part of the customer journey feels intentional, connected, and aligned with our broader strategy.

What you’ll do

Lead design initiatives that span multiple product areas, driving consistency, quality, and cohesion.

Partner with product teams to identify friction, gaps, and inconsistencies — and help define solutions that improve the overall jour